1Institute of Mental Health, National Clinical Research Center for Mental Disorders, Key Laboratory of Mental Health and Sixth Affiliated Hospital of Peking University,Beijing 100191, China 2Peking-Tsinghua Center for Life Sciences and PKU-IDG/McGovern Institute for Brain Research, Peking University,Beijing 100871, China 3Laboratory of Behavioral Neuroscience, Ningbo Addiction Research and Treatment Center, Medical School of Ningbo University, Ningbo 315010, China

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